TRAVERSE CITY, Mich. — A gray wolf relocated this fall from mainland Minnesota to Isle Royale National Park has died of unknown causes, officials said Tuesday, a minor setback in a multiyear plan to rebuild the predator species on the Lake Superior archipelago.

The 5-year-old male was among the first two wolves released at the park Sept. 26 . Staff biologists became concerned in late October when his radio tracking collar indicated he was no longer on the move, park spokeswoman Liz Valencia said.

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