Lee created The Fantastic Four, Spiderman, The Incredible Hulk and X-Men among others.
In its heyday, Marvel sold 50 million copies a year. Until he retired from editing in 1971, Lee wrote all the copy for Marvel's covers. He was also known for making a cameo in every Marvel film.
His characters brought the humans behind the heroes to life for the first time.
Hugh Jackman, who played Wolverine in the X-Men films, called Lee a "pioneering force in the superhero universe".
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